The evolutionary ancestry of arthropods dates back to the Cambrian period. The group is generally regarded as monophyletic, and many analyses support the placement of arthropods with cycloneuralians (or their constituent clades) in a superphylum Ecdysozoa. Overall, however, the basal relationships of animals are not yet well resolved. Likewise, the relationships between various arthropod groups are still actively debated. Today, arthropods contribute to the human food supply both directly as food, and more importantly, indirectly as pollinators of crops. Some species are known to spread severe disease to humans, livestock, and crops. (Full article...)
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Cimex lectularius
The Cimicidae are a family of small parasiticbugs that feed exclusively on the blood of warm-blooded animals. They are called cimicids or, loosely, bed bugs, though the latter term properly refers to the most well-known member of the family, Cimex lectularius, the common bed bug and its tropical relation Cimex hemipterus. The family contains over 100 species. Cimicids appeared in the fossil record in the Cretaceous period. When bats evolved in the Eocene, Cimicids switched hosts and now feed mainly on bats or birds. Members of the group have colonised humans on three occasions.
Cimicids usually feed on their host's blood every three to seven days, crawling away from the host and hiding while they digest the blood, which may take several days. This means that they specialise in vertebrate hosts that return regularly to particular sites to nest, roost or sleep. Birds and bats suit these specific requirements, as do humans now that they live in dwellings, and these are the main hosts used by the bugs. Most cimicids are able to go for long periods without feeding, over a year in some instances.
Cimicids are typically small, oval, flattened, wingless insects. They are stimulated to appear from their hiding places by cues such as a slight rise in the temperature of their surroundings. Among the family's distinctive characteristics are traumatic insemination, in which the male fertilises the eggs by piercing the female's abdominal wall with his intromittent organ. They also have distinctive paired structures called mycetomes inside their bodies, in which they harbour bacterial symbionts: these may help them to obtain nutrients they cannot get from blood. Although the insects may acquire viruses and other pathogens while feeding, these do not normally replicate inside the insect, and the infections are not transmitted to new hosts. (Full article...)
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Helicoverpa zea, commonly known as the corn earworm, is a species (formerly in the genus Heliothis) in the family Noctuidae. The larva of the mothHelicoverpa zea is a major agricultural pest. Since it is polyphagous (feeds on many different plants) during the larval stage, the species has been given many different common names, including the cotton bollworm and the tomato fruitworm. It also consumes a wide variety of other crops.
The species is widely distributed across the Americas with the exception of northern Canada and Alaska. It has become resistant to many pesticides, but can be controlled with integrated pest management techniques including deep ploughing, trap crops, chemical control using mineral oil, and biological controls.
The species migrates seasonally, at night, and can be carried downwind up to 400 km. Pupae can make use of diapause to wait out adverse environmental conditions, especially at high latitudes and in drought. (Full article...)
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The bush coconut, or bloodwood apple, is an Australian bush tucker food. It is an insect gall with both plant and animal components: an adult female scale insect and her offspring (of genus Cystococcus) live in a gall induced on a bloodwood eucalypt tree (Corymbia). Bush coconuts can vary from golf ball to tennis ball size. They have a hard and lumpy outer layer. The inner layer is a white flesh that contains the female insect and her offspring. There are three known species of Cystococcus responsible for forming the bush coconut: Cystococcus pomiformis, Cystococcus echiniformis and Cystococcus campanidorsalis. C. pomiformis is the most common species. The bush coconut is found in Western Australia, the Northern Territory, Queensland and New South Wales.
The bush coconut is picked from the host tree and cracked open to allow the flesh and scale insects to be eaten. Both have a high protein content and are used as a food source by humans and other animals. The name 'bush coconut' is derived from the white flesh of the inner layer, which is similar in appearance to that of a coconut, and the taste of the flesh has been said to have a coconut flavour. The bush coconut has been depicted in Indigenous Australian dreaming and used as inspiration in their artwork. (Full article...)
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Nephrops norvegicus, known variously as the Norway lobster, Dublin Bay prawn, shlobster (shrimp-lobster), langoustine (compare langostino) or shrimp, is a slim, coral colored lobster that grows up to 25 cm (10 in) long, and is "the most important commercial crustacean in Europe". It is now the only extant species in the genus Nephrops, after several other species were moved to the closely related genus Metanephrops. It lives in the north-eastern Atlantic Ocean, and parts of the Mediterranean Sea, but is absent from the Baltic Sea and Black Sea. Adults emerge from their burrows at night to feed on worms and fish. (Full article...)
Puzzled by the geographical distribution of wildlife and fossils he collected on the voyage, Darwin began detailed investigations and, in 1838, devised his theory of natural selection. Although he discussed his ideas with several naturalists, he needed time for extensive research, and his geological work had priority. He was writing up his theory in 1858 when Alfred Russel Wallace sent him an essay that described the same idea, prompting immediate joint submission of both their theories to the Linnean Society of London. Darwin's work established evolutionary descent with modification as the dominant scientific explanation of diversification in nature. In 1871, he examined human evolution and sexual selection in The Descent of Man, and Selection in Relation to Sex, followed by The Expression of the Emotions in Man and Animals (1872). His research on plants was published in a series of books, and in his final book, The Formation of Vegetable Mould, through the Actions of Worms (1881), he examined earthworms and their effect on soil. (Full article...)
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A. wedmannae holotype
Archimyrmex is an extinctgenus of ant in the formicid subfamily Myrmeciinae, described by palaeoentomologist Theodore Cockerell in 1923. The genus contains four described species, Archimyrmex rostratus, Archimyrmex piatnitzkyi, Archimyrmex smekali and Archimyrmex wedmannae. Archimyrmex is known from a group of Middle Eocene fossils which were found in North America, South America, and Europe. The genus was initially placed in the subfamily Ponerinae, but it was later placed in Myrmeciinae; it is now believed to be the ancestor of the extant primitive genus Myrmecia from Australia. Despite this, Archimyrmex is not a member to any tribe and is regarded as incertae sedis within Myrmeciinae. However, some authors believe Archimyrmex should be assigned as incertae sedis within Formicidae. These ants can be characterised by their large mandibles and body length, ranging from 13.2 to 30 mm (0.52 to 1.18 in). They also have long, thin legs and an elongated mesosoma (thorax) and petiole. (Full article...)
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Two wingless adults and a nymph
The black bean aphid (Aphis fabae) is a small black insect in the genus Aphis, with a broad, soft body, a member of the orderHemiptera. Other common names include blackfly, bean aphid, and beet leaf aphid. In the warmer months of the year, it is found in large numbers on the undersides of leaves and on the growing tips of host plants, including various agricultural crops and many wild and ornamental plants. Both winged and wingless forms exist, and at this time of year, they are all females. They suck sap from stems and leaves and cause distortion of the shoots, stunted plants, reduced yield, and spoiled crops. This aphid also acts as a vector for viruses that cause plant disease, and the honeydew it secretes may encourage the growth of sooty mould. It breeds profusely by live birth, but its numbers are kept in check, especially in the later part of the summer, by various predatory and parasitic insects. Ants feed on the honeydew it produces, and take active steps to remove predators. It is a widely distributed pest of agricultural crops and can be controlled by chemical or biological means. In the autumn, winged forms move to different host plants, where both males and females are produced. These mate and the females lay eggs which overwinter. (Full article...)
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Illustration of the carapace of E. batrachophthalmus
Eocarcinosoma is a genus of eurypterid, an extinct group of aquatic arthropods. The type and only species of Eocarcinosoma, E. batrachophthalmus, is known from deposits of Late Ordovician age in the United States. The generic name is derived from the related genus Carcinosoma, and the Greek eós (ἠώς) meaning 'dawn', referring to the earlier age of the genus compared to other carcinosomatid eurypterids.
Eocarcinosoma is known only from a single specimen, a well-preserved small prosoma (head). In life it would have been a very small eurypterid, given that the head measures just 2.05 centimetres (0.8 in) in length. The main distinguishing features from other carcinosomatids were the eyes being placed on the margins of the carapace (head plate) and the head being more triangular in shape than in its later relatives. Although initially believed to be an adult given the relatively small size of the eyes, some researchers believe Eocarcinosoma to represent a juvenile specimen of the eurypterid Megalograptus. (Full article...)
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Illustration of PWL2013/5224-LS, the type specimen of W. noctua
Wiedopterus is a genus of eurypterid, an extinct group of aquatic arthropods. The type and only species of Wiedopterus, W. noctua, is known from deposits of Early Devonian age in Germany. The generic name derives from the Wied river, which runs near the site of the initial discovery, and the species name, noctua, derives from Latinnoctua (owl) which refers to the superficial resemblance of the carapace (head plate) to an owl.
A small eurypterid, the type and only known specimen of Wiedopterus does not preserve the appendages or the underside of the body, making certain classification of the genus impossible. As such, it is formally classified as Eurypterinaincertae sedis (indicating uncertain placement within the Eurypterina suborder of eurypterids). Though not yet formally classified as such, Wiedopterus was suggested to probably belong to the Adelophthalmidae family on account of various features, including its general body outline, the position of its eyes, the first segment after its head being reduced in size as well as there being longitudinal ridges on the posterior segments of its body. (Full article...)
This woodwasp is an invasive species in many parts of the world, including Australia, New Zealand, North America, South America, and South Africa, where it has become a significant economic pest of pine trees. The wasp can attack a wide variety of pine species, although some species seem to be more susceptible than others, and stressed trees often are attacked.
Industrial melanism is widespread in the Lepidoptera (butterflies and moths), involving over 70 species such as Odontopera bidentata (scalloped hazel) and Lymantria monacha (dark arches), but the most studied is the evolution of the peppered moth, Biston betularia. It is also seen in a beetle, Adalia bipunctata (two-spot ladybird), where camouflage is not involved as the insect has conspicuous warning coloration, and in the seasnake Emydocephalus annulatus where the melanism may help in excretion of trace elements through sloughing of the skin. The rapid decline of melanism that has accompanied the reduction of pollution, in effect a natural experiment, makes natural selection for camouflage "the only credible explanation".
Other explanations for the observed correlation with industrial pollution have been proposed, including strengthening the immune system in a polluted environment, absorbing heat more rapidly when sunlight is reduced by air pollution, and the ability to excrete trace elements into melanic scales and feathers. (Full article...)
Pseudastacus (meaning "false Astacus", in comparison to the extant crayfish genus) is an extinct genus of decapod crustaceans that lived during the Jurassic period in Europe, and possibly the Cretaceous period in Lebanon. Many species have been assigned to it, though the placement of some species remains uncertain and others have been reassigned to different genera. Fossils attributable to this genus were first described by Georg zu Münster in 1839 under the name Bolina pustulosa, but the generic name was changed in 1861 after Albert Oppel noted that it was preoccupied. The genus has been placed into different families by numerous authors, historically being assigned to Nephropidae or Protastacidae. Currently, it is believed to be a member of Stenochiridae.
Reaching up to 6 cm (2.4 in) in total length, Pseudastacus was a small animal. Members of this genus have a crayfish-like build, possessing long antennae, a triangular rostrum and a frontmost pair of appendages enlarged into long and narrow pincers. Deep grooves are present on the carapace, which is around the same length as the abdomen. The surface of the carapace is usually uneven, with either small tubercles or pits. Sexual dimorphism is known in P. pustulosus, with the pincers of females being more elongated than those of the males. There is evidence of possible gregarious behavior in P. lemovices in the form of multiple individuals preserved alongside each other, possibly killed in a mass mortality event. With the oldest known record dating to the Sinemurian age of the Early Jurassic, and possible species surviving into the Cenomanian stage of the Late Cretaceous, Pseudastacus has a long temporal range and was a widespread taxon. Fossils of this animal were first found in the Solnhofen Limestone of Germany, but have also been recorded from France, England and Lebanon. All species in this genus lived in marine environments. (Full article...)
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Male M. campanulae
Megachile campanulae, known as the bellflower resin bee, is a species of bee in the family Megachilidae. Described in 1903, these solitary bees are native to eastern North America. Studies in 2013 placed them among the first insect species to use synthetic materials for making nests. They are considered mason bees, which is a common descriptor of bees in several families, including Megachilidae. Within the genus Megachile, frequently also referred to as leafcutter bees, M. campanulae is a member of the subgenus Chelostomoides, which do not construct nests from cut leaves, but rather from plant resins and other materials. Females lay eggs in nests constructed with individual cell compartments for each egg. Once hatched, the eggs progress through larval stages and subsequently will overwinter as pupae. The bees are susceptible to parasitism from several other bee species, which act as brood parasites. They are medium-sized bees and the female adults are typically larger than the males. They are important pollinators of numerous native plant species throughout their range. (Full article...)
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John S. Clark (21 March 1885 – 1 June 1956) was a Scottish-born Australian entomologist and myrmecologist known for his study of Australian ants. Born in Glasgow, he developed an interest in entomology at a young age. Clark first arrived in Australia in 1905 and originally worked for the state railways in Queensland. He developed an interest in ants shortly afterwards, collecting his first specimens in North Queensland. He married his first wife, Maggie Forbes in 1908, who bore four children, and died in 1935. He married his second wife, Phyllis Marjorie Claringbulls in 1939 and had two daughters with her. On her suicide in 1943, Clark sent his daughters to an orphanage.
In 1919, Clark worked as the assistant to the entomologist on probation in the Western Australian Department of Agriculture, but in 1920, he took on this position full-time. He started to publish his first articles about pest insects and ants from 1921; in 1926 he became an entomologist at the National Museum in Melbourne, remaining there for 20 years. Clark continued to publish more articles until he resigned from the National Museum of Victoria in 1944. Living in poverty, Clark lived a reclusive life, publishing his last book in 1951. He died on 1 June 1956 at the age of 71, survived by his six children. One of Clark's most notable achievements was describing Nothomyrmecia macrops, the most primitive living ant. Several ants have been named after him in recognition of his contributions. (Full article...)
Mecoptera (from the Greek: mecos = "long", ptera = "wings") is an order of insects in the superorder Holometabola with about six hundred species in nine families worldwide. Mecopterans are sometimes called scorpionflies after their largest family, Panorpidae, in which the males have enlarged genitals raised over the body that look similar to the stingers of scorpions, and long beaklike rostra. The Bittacidae, or hangingflies, are another prominent family and are known for their elaborate mating rituals, in which females choose mates based on the quality of gift prey offered to them by the males. A smaller group is the snow scorpionflies, family Boreidae, adults of which are sometimes seen walking on snowfields. In contrast, the majority of species in the order inhabit moist environments in tropical locations.
The Mecoptera are closely related to the Siphonaptera (fleas), and a little more distantly to the Diptera (true flies). They are somewhat fly-like in appearance, being small to medium-sized insects with long slender bodies and narrow membranous wings. Most breed in moist environments such as leaf litter or moss, and the eggs may not hatch until the wet season arrives. The larvae are caterpillar-like and mostly feed on vegetable matter, and the non-feeding pupae may pass through a diapause until weather conditions are favorable.
Early Mecoptera may have played an important role in pollinating extinct species of gymnosperms before the evolution of other insect pollinators such as bees. Adults of modern species are overwhelmingly predators or consumers of dead organisms. In a few areas, some species are the first insects to arrive at a cadaver, making them useful in forensic entomology. (Full article...)
... that the family placement for the fossil moth genus Dominickus was not noticed until entomologistNorman Tindale was looking at pictures of modern moths from Australia?
Image 7 This fully-grown robber crab has tough fabric forming its joints, delicate biomineralized cuticle over its sensory antennae, optic-quality over its eyes, and strong, calcite-reinforced chitin armouring its body and legs; its pincers can break into coconuts (from Arthropod exoskeleton)
Image 9In honeypot antrepletes, the abdomens of the workers that hold the sugar solution grow vastly, but only the unsclerotised cuticle can stretch, leaving the unstretched sclerites as dark islands on the clear abdomen (from Arthropod exoskeleton)
Image 14Ghost crab, showing a variety of integument types in its exoskeleton, with transparent biomineralization over the eyes, strong biomineralization over the pincers, and tough chitin fabric in the joints and the bristles on the legs (from Arthropod exoskeleton)
Image 15Body structure of a typical crustacean – krill (from Crustacean)
Image 16 Honeybee larvae have flexible but delicate unsclerotised cuticles. (from Arthropod exoskeleton)
Image 18This Zoea-stage larva is hardly recognisable as a crab, but each time it sheds its cuticle it remodels itself, eventually taking on its final crab form (from Arthropod exoskeleton)
Image 19Decapods, from Ernst Haeckel's 1904 work Kunstformen der Natur (from Crustacean)
Image 23Formation of anterior segments across arthropod taxa based on gene expression and neuroanatomical observations, Note the chelicera(Ch) and chelifore(Chf) arose from somite 1 and thus correspond to the first antenna(An/An1) of other arthropods. (from Chelicerata)
Image 25The house centipedeScutigera coleoptrata has rigid sclerites on each body segment. Supple chitin holds the sclerites together and connects the segments flexibly. Similar chitin connects the joints in the legs. Sclerotised tubular leg segments house the leg muscles, their nerves and attachments, leaving room for the passage of blood to and from the hemocoel (from Arthropod exoskeleton)
Image 26Crab larva barely recognisable as a crab, radically changes its form when it undergoes ecdysis as it matures (from Arthropod exoskeleton)
Image 27The fangs in spiders' chelicerae are so sclerotised as to be greatly hardened and darkened (from Arthropod exoskeleton)
Image 28Some of the various hypotheses of myriapod phylogeny. Morphological studies (trees a and b) support a sister grouping of Diplopoda and Pauropoda, while studies of DNA or amino acid similarities suggest a variety of different relationships, including the relationship of Pauropoda and Symphyla in tree c. (from Myriapoda)
Image 32Mature queen of a termite colony, showing how the unsclerotised cuticle stretches between the dark sclerites that failed to stretch as the abdomen grew to accommodate her ovaries (from Arthropod exoskeleton)
The Ozyptila praticola species of crab spider is found throughout Europe and the Middle East. They do not build webs to trap prey, but are active hunters. Crab spiders are so named because of their first two pairs of legs, which are held out to the side giving them a crab-like appearance. Also, like crabs, these spiders move sideways and backwards more easily than forwards.
The light blue soldier crab (Mictyris longicarpus) inhabits beaches in the Indo-Pacific region. Soldier crabs filter sand or mud for microorganisms. They congregate during the low tide, and bury themselves in a corkscrew pattern during high tide, or whenever they are threatened.
Two Melangyna viridiceps (called Common Hoverflies in Australia) mating in mid-air. The male, which can be identified by the eyes meeting at the top of its head, is on top. The term "hoverfly" refers to about 6,000 species of flying insects in the family Syrphidae. They are often seen hovering at flowers and are important pollinators.
A fishing spider with two of its legs missing. Most likely some predator (a bird, or given its habitat a large fish or frog) grasped the spider by the missing limbs which were jettisoned by the spider in response, a process known as autotomy.
The orb-weaver spiders (family Araneidae) are the familiar builders of spiral wheel-shaped webs often found in gardens, fields and forests. The family is a large one, including over 2800 species in over 160 genera worldwide, making it the third largest known (behind Salticidae and Linyphiidae). The web has always been thought of as an engineering marvel.
A macro shot of the head of a dragonfly, focusing on its compound eyes. Dragonfly eyes have up to 30,000 facets; each one is a separate light-sensing organ or ommatidium, arranged to give nearly a 360° field of vision.
Argiope trifasciata, the banded garden or banded orb-weaving spider, is a species of arachnid in the family Araneidae. It is native to North and South America but has spread to other parts of the world. This ventral view of a female A. trifasciata shows her in the centre of her web, which can reach a diameter of 60 cm (24 in). The function of the zig-zag web decorations is unclear, but they may serve to make the spider appear larger or to act as a warning sign.
The Adonis blue (Polyommatus bellargus) is a butterfly in the family Lycaenidae, found in Europe and parts of West Asia. This photograph, taken in a chalk meadow at Yoesden Bank in Buckinghamshire, England, shows the underside of the folded wings of a male Adonis blue; the upper side is a bright, sky-blue.
The Forest scorpion (Cercophonius squama) is a scorpion native to southeastern Australia and Tasmania. The body is 25 to 40 millimetres (0.98 to 1.57 in) long, and coloured creamy yellow to orange brown with dark brown variegations. The legs are yellow with some dark brown pigment.
A mole cricket, an insect belonging to the Gryllotalpidaefamily. Mole crickets are common insects, found on every continent except Antarctica, but because they are nocturnal and spend nearly all their lives underground in extensive tunnel systems, they are rarely seen. This specimen is likely to be Gryllotalpa brachyptera and is about 3.5 cm (1.4 in.) in size.
A nest of the tropical Asian giant honey bee, which consists of a single exposed honeycomb, an array of densely packed hexagonal cells made of beeswax. Honeycombs store food (honey and pollen) and house the "brood" (eggs, larvae, and pupae).
Purana tigrina is a species of cicada found in Southeast Asia. This adult male was photographed in Kadavoor, Kerala, in southern India, and is about one inch (25 mm) in length. The mouthparts are adapted to piercing plant tissues and sucking sap; the male abdomen houses the tymbal, an organ used in the production of song, while the female abdomen is tipped by a large, saw-edged ovipositor.
Xylotrupes socrates (Siamese rhinoceros beetle, or "fighting beetle"), male, on a banana leaf. This scarab beetle is particularly known for its role in insect fighting in Northern Laos and Thailand.
The monarch butterfly (Danaus plexippus) is one of the best-known species of butterfly. Its wings feature an easily recognizable orange and black pattern, with a wingspan of 8.9–10.2 centimetres (3½–4 in).
The Painted Lady (Vanessa cardui) is a well-known colourful butterfly, found on every continent except Antarctica. It occurs in any temperate zone, including mountains in the tropics. The species is resident only in warmer areas, but migrates in spring, and sometimes again in autumn.
Danaus genutia, the common tiger or striped tiger, is a species of brush-footed butterfly found in Sri Lanka, India, Myanmar, south-eastern Asia and Australia. It prefers areas of moderate to heavy rainfall, and typical habitats include scrubby jungle, deciduous forests and fallow land near habitations. The insect sequesters toxins from plants, and advertises its unpalatability by having prominent markings and striking colour patterns. This adult male common tiger, of the subspecies D. g. genutia, was photographed in Kerala, India.
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